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David H. Ward

Title: Research Wildlife Biologist
Address: 4210 University Dr., Anchorage, AK 99508-4626
Phone: (907) 786-7097
Fax: (907) 786-7021

Image of David H. Ward

Education and/or Training

M.S.1983University of Oregon, Charleston, ORBiology with Marine Emphasis
B.A.1974Hartwick College, Oneonta, NYBiology

Areas of Specialization and/or Research Interests

Population ecology, wintering and migration ecology of arctic breeding waterbirds; impacts of disturbance on migratory birds; mapping and remote sensing of seagrasses.

Professional Experience

1997 - PresentResearch Wildlife Biologist, USGS, Alaska Science Center, Anchorage, Alaska
1994 - 1997Research Wildlife Biologist, NBS, Alaska Biological Science Center, Anchorage, Alaska
1985 - 1994Research Wildlife Biologist, USFWS, Alaska Fish and Wildlife Research Center, Anchorage, Alaska

Professional Activities and/or Memberships

Cooper Ornithological Society
American Birding Association
Association of Field Ornithologists
American Ornithological Union
The Wildlife Society
Wilson Ornithological Society

Websites of Interest

Wintering Ecology of Surf Scoters in Southeast Alaska

Pacific Brant Opting Out of Migration


Total pubs: 65

Hupp, J. W., D. H. Ward, K. R. Hogrefe, J. G. Sedinger, P. D. Martin, A. Stickney, and T. Obritschkewitsch. 2017. Growth of black brant and snow goose goslings in northern Alaska: implications for increasing goose populations. Journal of Wildlife Management 81(5):846-857. doi:10.1002/jwmg.21246 [Details] [Full Publication]

Leach, A. G., D. H. Ward, J. G. Sedinger, M. S. Lindberg, W. S. Boyd, J. W. Hupp, and R. J. Ritchie. 2017. Declining survival of black brant from subarctic and arctic breeding areas. Journal of Wildlife Management 81(7):1210-1218. doi:10.1002/jwmg.21284 [Details] [Full Publication]

Uher-Koch, B. D., D. Esler, S. A. Iverson, D. H. Ward, S. Boyd, M. Kirk, T. L. Lewis, C. S. VanStratt, K. M. Brodhead, J. W. Hupp, and J. A. Schmutz. 2016. Interacting effects of latitude, mass, age, and sex on winter survival of Surf Scoters (Melanitta perspicillata): implications for differential migration. Canadian Journal of Zoology 94(3):233-241. doi:10.1139/cjz-2015-0107 [Details] [Full Publication]

Talbot, S. L., G. K. Sage, J. R. Rearick, M. C. Fowler, R. Muñiz-Salazar, B. Baibak, S. Wyllie-Echeverria, A. Cabello-Pasini, and D. H. Ward. 2016. The structure of genetic diversity in eelgrass (Zostera marina L.) along the North Pacific and Bering Sea coasts of Alaska. PLoS One. doi:10.1371/journal.pone.0152701 [Details] [Full Publication]

Weiser, E. L., R. B. Lanctot, S. C. Brown, J. A. Alves, P. F. Battley, R. Bentzen, J. Bêty, M. A. Bishop, M. Boldenow, L. Bollache, B. Casler, M. Christie, J. T. Coleman, J. R. Conklin, W. B. English, H. R. Gates, O. Gilg, M-A. Giroux, K. Gosbell, C. J. Hassell, J. Helmericks, A. Johnson, B. Katrínardóttir, K. Koivula, E. Kwon, J-F. Lamarre, J. Lang, D. B. Lank, N. Lecomte, J. R. Liebezeit, V. Loverti, L. McKinnon, C. Minton, D. Mizrahi, E. Nol, V-M. Pakanen, J. Perz, R. Porter, J. Rausch, J. Reneerkens, N. Rönkä, S. Saalfeld, N. R. Senner, B. Sittler, P. A. Smith, K. M. Sowl, A. Taylor, D. H. Ward, S. Yezerinac, and B. K. Sandercock. 2016. Effects of geolocators on hatching success, return rates, breeding movements, and change in body mass in 16 species of Arctic-breeding shorebirds. Movement Ecology 4(12). doi:10.1186/s40462-016-0077-6 [Details] [Full Publication]

Ward, D. H., J. Helmericks, J. W. Hupp, L. McManus, M. Budde, D. C. Douglas, and K. D. Tape. 2015. Multi-decadal trends in spring arrival of avian migrants to the central Arctic coast of Alaska: effects of environmental and ecological factors. Journal of Avian Biology 47(2):197-207. doi:10.1111/jav.00774 [Details] [Full Publication]

Hupp, J. W., D. H. Ward, M. E. Whalen, and J. M. Pearce. 2015. Changing Arctic ecosystems—What is causing the rapid increase of Snow Geese in northern Alaska?. USGS Fact Sheet 2015-3062, 2 p. doi:10.3133/fs20153062 [Details] [Full Publication]

Stillman, R. A., K. A. Wood, W. Gilkerson, E. Elkinton, J. M. Black, and D. H. Ward. 2015. Predicting effects of environmental change on a migratory herbivore. Ecosphere 6:art114. doi:10.1890/ES14-00455.1 [Details] [Full Publication]

Petrich, C., A. C. Tivy, and D. H. Ward. 2014. Reconstruction of historic sea ice conditions in a sub-Arctic lagoon. Cold Regions Science and Technology 98:55-62. doi:10.1016/j.coldregions.2013.10.011 [Details] [Full Publication]

Ramey, A. M., J. A. Reed, J. A. Schmutz, T. F. Fondell, B. W. Meixell, J. W. Hupp, D. H. Ward, J. Terenzi, and C. R. Ely. 2014. Prevalence, transmission, and genetic diversity of blood parasites infecting tundra-nesting geese in Alaska. Canadian Journal of Zoology 92(8):699-706. doi:10.1139/cjz-2014-0041 [Details] [Full Publication]

Liebezeit, J. R., K. E. B. Gurney, M. Budde, S. Zack, and D. H. Ward. 2014. Phenological adjustment in arctic bird species: relative importance of snow melt and ecological factors. Polar Biology 37(9):1309-1320. doi:10.1007/s00300-014-1522-x [Details] [Full Publication]

Tape, K. D., J. M. Pearce, D. Walworth, B. W. Meixell, T. F. Fondell, D. D. Gustine, P. L. Flint, J. W. Hupp, J. A. Schmutz, and D. H. Ward. 2014. Historical and contemporary imagery to assess ecosystem change on the Arctic coastal plain of northern Alaska (ver. 1.1, June 2015). USGS Open-File Report 2014-1140, 22 p. doi:10.3133/ofr20141140 [Details] [Full Publication]

Hogrefe, K. R., D. H. Ward, T. F. Donnelly, and N. C. Dau. 2014. Establishing a Baseline for Regional Scale Monitoring of Eelgrass (Zostera marina) Habitat on the Lower Alaska Peninsula. Remote Sensing 6(12):12447-12477. doi:10.3390/rs61212447 [Details] [Full Publication]

Boyd, W. S., D. H. Ward, D. K. Kraege, and A. A. Gerick. 2013. Migration patterns of Western High Arctic (Grey-belly) Brant Branta bernicla. Wildfowl Special Issue 3:3-25. [Details] [Full Publication]

Lewis, T. L., D. H. Ward, J. S. Sedinger, A. Reed, and D. V. Derksen. 2013. Brant (Branta bernicla), The Birds of North America Online (A. Poole, Ed.). Ithaca: Cornell Lab of Ornithology. Birds of North America Online. Retrieved from [Details] [Full Publication]

Martinez, I., R. Carmona, D. H. Ward, and G. D. Danemann. 2013. Habitat use patterns of the Black Brant Branta bernicla nigricans (Anseriformes: Anatidae) in natural and artificial areas of Guerrero Negro, Baja California Sur, Mexico. International Journal of Tropical Biology and Conservation 61(2):927-935. [Details]

Nicolai, C. A., J. S. Sedinger, D. H. Ward, and W. S. Boyd. 2012. Mate loss affects survival but not breeding in Black Brant geese. Behavioral Ecology 23(3):643-648. doi:10.1093/beheco/ars009 [Details] [Full Publication]

Lok, E. K., D. Esler, J. Y. Takekawa, S. W. De La Cruz, W. S. Boyd, D. R. Nysewander, J. R. Evenson, and D. H. Ward. 2012. Spatiotemporal associations between Pacific herring spawn and Surf Scoter spring migration: evaluating a 'silver wave' hypothesis. Marine Ecology Progress Series 457:139-150. [Details]

Shaughnessy, F. J., W. Gilkerson, J. M. Black, D. H. Ward, and M. Petrie. 2012. Predicted eelgrass response to sea level rise and its availability to foraging Black Brant in Pacific coast estuaries. Ecological Applications 22(6):1743-1761. doi:10.1890/11-1083.1 [Details] [Full Publication]

Schamber, J. L., J. S. Sedinger, and D. H. Ward. 2012. Carry-over effects of winter location contribute to variation in timing of nest initiation and clutch size in Black Brant (Branta bernicla nigricans). The Auk 129(2):205-210. doi:10.1525/auk.2012.11249 [Details] [Full Publication]

Pearce, J. M., A. R. DeGange, P. L. Flint, T. F. Fondell, D. D. Gustine, L. E. Holland-Bartels, A. G. Hope, J. W. Hupp, J. C. Koch, J. A. Schmutz, S. L. Talbot, D. H. Ward, and M. E. Whalen. 2012. Changing Arctic Ecosystems - Measuring and forecasting the response of Alaska's terrestrial ecosystem to a warming climate. USGS Fact Sheet 2012-3144, 4 p. [Details] [Full Publication]

Anderson, E. M., D. Esler, W. S. Boyd, J. R. Evenson, D. R. Nysewander, D. H. Ward, R. D. Dickson, B. D. Uher-Koch, C. S. VanStratt, and J. W. Hupp. 2012. Predation rates, timing, and predator composition for scoters (Melanitta spp.) in marine habitats. Canadian Journal of Zoology 90(1):42-50. doi:10.1139/z11-110 [Details] [Full Publication]

Takekawa, J. Y., S. W. De La Cruz, M. T. Wilson, E. C. Palm, J. Lee, D. R. Nysewander, J. R. Evenson, D. Esler, W. S. Boyd, and D. H. Ward. 2011. Breeding synchrony, sympatry, and nesting areas of Pacific coast surf scoters (Melanitta perspicillata). Pages 41-64 in J.V. Wells, (ed.). Boreal birds of North America: a hemispheric view of their conservation links and significance. Studies in Avian Biology (no. 41). University of California Press, Berkeley, CA, USA. [Details]

Sedinger, J. G., J. L. Schamber, D. H. Ward, C. A. Nicolai, and B. Conant. 2011. Carryover effects associated with winter location affect fitness, social status, and population dynamics in a long-distant migrant. American Naturalist 178(5):E110-E123. doi:10.1086/662165 [Details] [Full Publication]

Lok, E. K., D. Esler, J. Y. Takekawa, S. W. De La Cruz, W. S. Boyd, D. R. Nysewander, J. R. Ball, and D. H. Ward. 2011. Stopover habitats of spring migrating Surf Scoters in southeast Alaska. Journal of Wildlife Management 75:92-100. [Details]

Black, J. M., D. E. Lee, and D. H. Ward. 2010. Foraging home ranges of Black Brant Branta bernicla nigricans during spring stopover at Humboldt Bay, California, USA. Wildfowl 60:85-94. [Details] [Full Publication]

Ward, D. H., C. P. Dau, T. L. Tibbitts, J. G. Sedinger, B. A. Anderson, and J. E. Hines. 2009. Change in abundance of Pacific Brant wintering in Alaska: Evidence of a climate warming effect?. Arctic 62(3):301-311. [Details] [Full Publication]

De La Cruz, S. W., J. Y. Takekawa, M. T. Wilson, D. R. Nysewander, J. R. Evenson, D. Esler, W. S. Boyd, and D. H. Ward. 2009. Surf Scoter (Melanitta perspicillata) spring migration routes and chronology: A synthesis of Pacific coast studies. Canadian Journal of Zoology 87:1069-1086. doi:10.1139/Z09-099 [Details] [Full Publication]

Sedinger, J. S., N. D. Chelgren, D. H. Ward, and M. S. Lindberg. 2008. Fidelity and breeding probability related to population density and individual quality in Black Brant geese (Branta bernicla nigricans). Journal of Animal Ecology 77(4):702-712. doi:10.1111/j.1365-2656.2008.01403.x [Details] [Full Publication]

Lindberg, M. S., D. H. Ward, T. L. Tibbitts, and J. Roser. 2007. Winter movement dynamics of a migratory bird species, Black Brant. Journal of Wildlife Management 71(2):534-540. [Details]

Schamber, J. L., J. S. Sedinger, D. H. Ward, and K. R. Hagemeier. 2007. Latitudinal variation in population structure of wintering Pacific black brant. Journal of Field Ornithology 78(1):74-82. [Details]

Mason, D. D., P. S. Barboza, and D. H. Ward. 2007. Mass dynamics of wintering brant: body, adipose tissue, organ, and muscle mass vary with location. Canadian Journal of Zoology 85(6):728-736. doi:10.1139/Z07-050 [Details] [Full Publication]

Talbot, S. L., S. Wyllie-Echeverria, D. H. Ward, J. R. Rearick, G. K. Sage, B. Chesney, and R. C. Phillips. 2006. Genetic characterization of Zostera asiatica on the Pacific Coast of North America. Aquatic Botany 85(3):169-176. doi:10.1016/j.aquabot.2006.03.011 [Details] [Full Publication]

Muñiz-Salazar, R., S. L. Talbot, G. K. Sage, D. H. Ward, and A. Cabello-Pasini. 2006. Genetic structure of eelgrass Zostera marina meadows in an embayment with restricted water flow. Marine Ecology Progress Series 309:107-116. [Details] [Full Publication]

Sedinger, J. S., D. H. Ward, J. L. Schamber, W. I. Butler, W. D. Eldridge, B. Conant, J. F. Voelzer, N. D. Chelgren, and M. P. Herzog. 2006. Effects of El Niño on distribution and reproductive performance of Black Brant. Ecology 87(1):151-159. [Details]

Mason, D. D., P. S. Barboza, and D. H. Ward. 2006. Nutritional condition of Pacific Black Brant wintering at the extremes of their range. Condor 108(3):678-690. [Details] [Full Publication]

Muñiz-Salazar, R., S. L. Talbot, G. K. Sage, D. H. Ward, and A. Cabello-Pasini. 2005. Population genetic structure of annual and perennial populations of Zostera marina L. along the Pacific coast of Baja California and the Gulf of California. Molecular Ecology 14(3):711-722. doi:10.1111/j.1365-294X.2005.02454.x [Details] [Full Publication]

Ward, D. H., A. Reed, J. S. Sedinger, J. M. Black, D. V. Derksen, and P. M. Castelli. 2005. North American Brant: Effects of changes in habitat and climate on population dynamics. Global Change Biology 11:869-880. doi:10.1111/j.1365-2486.2005.00942.x [Details] [Full Publication]

Cabello-Pasini, A., R. Muñiz-Salazar, and D. H. Ward. 2004. Biochemical characterization of the eelgrass Zostera marina at its southern distribution limit in the North Pacific. Ciencias Marinas 30(1a):21-24. [Details] [Full Publication]

Ward, D. H., T. L. Tibbitts, A. Morton, E. Carrera-Gonzales, and R. Kempka. 2004. Use of digital multispectral videography to assess seagrass distribution in San Quintín, Baja California, Mexico. Ciencias Marinas 30(1A):47-60. [Details] [Full Publication]

Ackerman, J. T., J. Y. Takekawa, K. L. Kruse, D. L. Orthmeyer, J. L. Yee, C. R. Ely, D. H. Ward, K. S. Bollinger, and D. M. Mulcahy. 2004. Using radiotelemetry to monitor cardiac response of free living Tule greater white-fronted geese (Anser albifrons elgasi) to human disturbance. Wilson Bulletin 116(2):146-151. [Details] [Full Publication]

Ward, D. H., J. A. Schmutz, J. S. Sedinger, K. S. Bollinger, P. D. Martin, and B. A. Anderson. 2004. Temporal and geographic variation in survival of juvenile Black Brant. Condor 106:263-274. [Details] [Full Publication]

Sedinger, J. S., M. P. Herzog, and D. H. Ward. 2004. Early Environment and Recruitment of Black Brant (Brant Bernicla Nigricans) Into the Breeding Population. The Auk 121(1):68-73. [Details]

Cabello-Pasini, A., R. Muñiz-Salazar, and D. H. Ward. 2003. Annual variation of biomass and photosynthesis in Zostera marina L. along the Pacific Coast of Baja California, Mexico. Aquatic Botany 76:31-47. [Details]

Ward, D. H. and S. Ibarra-Obando. 2003. Seagrasses of the Pacific Coast of North America. Pages 218 in E. Green and F. Short, (eds.). World atlas of seagrasses: present and future conservation. University of California Press, Berkeley, CA, USA. [Details]

Ward, D. H., A. Morton, T. L. Tibbitts, D. C. Douglas, and E. Carrera-Gonzales. 2003. Long-term change in eelgrass distribution at Bahía San Quintín Baja California, Mexico, using satellite imagery. Estuaries 26(6):1529-1539. doi:10.1007/BF02803661 [Details] [Full Publication]

Ward, D. H., A. Morton, T. L. Tibbitts, S. Ibarra-Obando, and D. C. Douglas. 2003. Recent trends in eelgrass, Zostera marina, distribution and abundance along the west coast of Baja California, Mexico. Pages 139 in Proceedings of the 5th International Seagrass Biology Workshop, Ensenada, Baja California, October 2002. Gulf of Mexico Science 21 [Details]

Sedinger, J. S., N. D. Chelgren, M. S. Lindberg, T. Obritschkewitch, M. Kirk, P. D. Martin, and B. A. Anderson, and D. H. Ward. 2002. Life-history implications of large-scale variation in adult survival of black brant (Branta bernicla nigricans). The Auk 119(2):510-515. [Details] [Full Publication]

Ward, D. H., R. A. Stehn, and D. V. Derksen. 2000. Response of Geese to Aircraft Disturbances. Terra Borealis 2:52-55. [Details]

Ely, C. R., D. H. Ward, and K. S. Bollinger. 1999. Behavioral correlates of heart rates of free-living greater white-fronted geese. Condor 101:390-395. [Details] [Full Publication]

Ward, D. H., R. A. Stehn, W. P. Erickson, and D. V. Derksen. 1999. Response of fall-staging brant and Canada geese to aircraft overflights in southwestern Alaska. Journal of Wildlife Management 63:373-381. [Details]

Erickson, W. P., T. G. Nick, and D. H. Ward. 1998. Investigating flight response of Pacific brant to helicopters at Izembek Lagoon, Alaska, by using logistic regression. Society for Industrial and Applied Mathematics:155-170. [Details]

Reed, A., D. H. Ward, D. V. Derksen, and J. S. Sedinger. 1998. Brant (Branta bernicla). No. 337 in A. Poole and F. Gill, (eds.). The Birds of North America, Philadelphia, PA, USA. [Details]

Sedinger, J. S., M. S. Lindberg, E. A. Rexstad, N. D. Chelgren, and D. H. Ward. 1997. Testing for handling bias in survival estimation for Black Brant. Journal of Wildlife Management 61:782-791. [Details]

Ward, D. H., C. J. Markon, and D. C. Douglas. 1997. Distribution and stability of eelgrass beds at Izembek Lagoon, Alaska. Aquatic Botany 58:229-240. doi:10.1016/S0304-3770(97)00037-5 [Details] [Full Publication]

Ward, D. H., E. A. Rexstad, J. S. Sedinger, M. S. Lindberg, and N. K. Dawe. 1997. Seasonal and annual survival of adult Pacific brant. Journal of Wildlife Management 61:773-781. [Details]

Derksen, D. V., K. S. Bollinger, D. H. Ward, J. S. Sedinger, and Y. Miyabayashi. 1996. Black brant from Alaska staging and wintering in Japan. Condor 98:653-657. doi:10.2307/1369583 [Details] [Full Publication]

Schmutz, J. A., D. H. Ward, and et al. 1995. Survival estimation and the effects of dependency among animals. Journal of Applied Statistics 22(5&6):673-681. [Details]

Ward, D. H. and P. L. Flint. 1995. Effects of harness-attached transmitters on premigration and reproduction of Brant. Journal of Wildlife Management 59:39-46. [Details]

Sedinger, J. S., D. H. Ward, R. M. Anthony, D. V. Derksen, C. J. Lensink, K. S. Bollinger, and N. K. Dawe. 1994. Management of Pacific brant: population structure and conservation issues. Transactions of the North American Wildlife and Natural Resources Conference 59:50-62. [Details]

Ward, D. H., R. A. Stehn, and D. V. Derksen. 1994. Response of staging brant to disturbance at Izembek Lagoon, Alaska. Wildlife Society Bulletin 22:220-228. [Details]

Sedinger, J. S., C. J. Lensink, D. H. Ward, R. M. Anthony, M. L. Wege, and G. V. Byrd. 1993. Current status and recent dynamics of the black brant breeding population. Wildfowl 44:49-59. [Details]

Derksen, D. V. and D. H. Ward. 1993. Life history strategies and habitat needs of the black brant. US Fish and Wildlife Leaflet 13.1.15. [Details]

Ward, D. H., D. V. Derksen, S. P. Kharitonov, M. Stishov, I. Stirling, V. V. Baranyuk. 1993. Status of Pacific black brant Branta bernicla nigricans on Wrangel Island, Russian Federation. Wildfowl 44:39-48. [Details]

Reed, A., R. A. Stehn, and D. H. Ward. 1989. Autumn use of Izembek Lagoon, Alaska, by brant from different breeding areas. Journal of Wildlife Management 53:720-725. [Details]

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