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Alaska Science Center

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George Kevin Sage

Title: Geneticist
Address: 4210 University Dr., Anchorage, AK 99508-4626
Phone: (907) 786-7189
Fax: (907) 786-7021

Education and/or Training

M.S.1993University of Montana, Missoula, MTZoology
B.S.1985University of Montana, Missoula, MTWildlife Biology

Areas of Specialization and/or Research Interests

Conservation genetics of North American salmon and trout.

Professional Experience

1996 - PresentResearch Geneticist, USGS, Alaska Science Center, Anchorage, Alaska
1989 -1995Research Assistant, University of Montana, Missoula, Montana
1986 - 1988Fisheries Field Worker, Montana Department of Fish, Wildlife, and Parks, Kalispell, Montana

Professional Activities and/or Memberships

American Fisheries Society
Alaska Chapter American Fisheries Society
Montana Chapter American Fisheries Society


Total pubs: 52

Sonsthagen, S. A., J. C. Williams, G. S. Drew, C. M. White, G. K. Sage, and S. L. Talbot. 2017. Legacy or colonization? Posteruption establishment of peregrine falcons (Falco peregrinus) on a volcanically active subarctic island. Ecology and Evolution 7(1):107-114. doi:10.1002/ece3.2631 [Details] [Full Publication]

Gravley, M. C., G. K. Sage, J. A. Schmutz, and S. L. Talbot. 2017. Development of microsatellite loci exhibiting reverse ascertainment bias and a sexing marker for use in Emperor Geese (Chen canagica). Avian Biology Research 10(4):201-210. doi:10.3184/175815617X14969254461396 [Details] [Full Publication]

Scribner, K. T., C. Soiseth, J. J. McGuire, G. K. Sage, L. K. Thorsteinson, J. L. Nielsen, and E. E. Knudsen. 2017. Genetic assessment of the effects of streamscape succession on coho salmon Oncorhynchus kisutch colonization in recently deglaciated streams. Journal of Fish Biology 91(1):195-218. doi:10.1111/jfb.13337 [Details] [Full Publication]

Talbot, S., G. K. Sage, S. A. Sonsthagen, M. C. Gravley, T. Swem, J. C. Williams, J. Longmire, S. Ambrose, M. J. Flamme, S. B. Lewis, L. Phillips, C. Anderson, and C. M. White. 2017. Intraspecific evolutionary relationships among peregrine falcons in western North American high latitudes. PLoS One 12(11):e0188185. doi:10.1371/journal.pone.0188185 [Details] [Full Publication]

Hollenbeck, N., D. Scheel, M. C. Gravley, G. K. Sage, R. K. Toussaint, and S. L. Talbot. 2017. Use of swabs for sampling epithelial cells for molecular genetics analyses in Enteroctopus. American Malacological Bulletin 35(2):145-157. doi:10.4003/006.035.0207 [Details] [Full Publication]

Talbot, S. L., G. K. Sage, J. R. Rearick, M. C. Fowler, R. Muñiz-Salazar, B. Baibak, S. Wyllie-Echeverria, A. Cabello-Pasini, and D. H. Ward. 2016. The structure of genetic diversity in eelgrass (Zostera marina L.) along the North Pacific and Bering Sea coasts of Alaska. PLoS One. doi:10.1371/journal.pone.0152701 [Details] [Full Publication]

Roffler, G. H., M. K. Schwartz, K. L. Pilgrim, S. L. Talbot, G. K. Sage, L. G. Adams, and G. Luikart. 2016. Identification of landscape features influencing gene flow: How useful are habitat selection models?. Evolutionary Applications 9(6):805-817. doi:10.1111/eva.12389 [Details] [Full Publication]

Haughey, C. L., G. K. Sage, G. R. DeGange, S. A. Sonsthagen, and S. L. Talbot. 2016. Development of novel microsatellite markers for the Northern Goshawk (Accipiter gentilis) and their utility in cross-species amplification. Avian Biology Research 9(3):195-199. doi:10.3184/175815516X14667737479433 [Details] [Full Publication]

Peacock, E., S. A. Sonsthagen, M. E. Obbard, A. N. Boltunov, E. V. Regehr, N. Ovsyanikov, J. Aars, S. N. Atkinson, G. K. Sage, A. G. Hope, E. Zeyl, L. Bachmann, D. Ehrich, K. T. Scribner, S. C. Amstrup, S. E. Belikov, E. W. Born, A. E. Derocher, I. Stirling, M. K. Taylor, Ø. Wiig, D. Paetkau, and S. L. Talbot. 2015. Implications of the circumpolar genetic structure of polar bears for their conservation in a rapidly warming Arctic. PLoS One 10(1): e112021. doi:10.1371/journal.pone.0112021 [Details] [Full Publication]

Sonsthagen, S. A., K. Fales, C. V. Jay, G. K. Sage, and S. L. Talbot. 2014. Spatial variation and low diversity in the major histocompatibility complex in walrus (Odobenus rosmarus). Polar Biology 37(4):497-506. doi:10.1007/s00300-014-1450-9 [Details] [Full Publication]

Farley, S. D., S. L. Talbot, G. K. Sage, R. Sinnott, and J. Coltrane. 2014. Use of DNA from bite marks to determine species and individuals animals that attack humans. Wildlife Society Bulletin 38(2):370-376. doi:10.1002/wsb.391 [Details] [Full Publication]

Roffler, G. H., S. L. Talbot, G. Luikart, G. K. Sage, K. L. Pilgrim, L. G. Adams, and M. K. Schwartz. 2014. Lack of sex-biased dispersal promotes fine-scale genetic structure in alpine ungulates. Conservation Genetics 15(4):837-851. doi:10.1007/s10592-014-0583-2 [Details] [Full Publication]

White, C. M., S. A. Sonsthagen, G. K. Sage, C. M. Anderson, and S. L. Talbot. 2013. Genetic relationships among some subspecies of the Peregrine Falcon (Falco peregrinus L.), inferred from mitochondrial DNA control-region sequences. The Auk 130(1):78-87. doi:10.1525/auk.2012.11173 [Details] [Full Publication]

Sonsthagen, S. A., G. K. Sage, M. C. Fowler, A. G. Hope, J. A. Cook, and S. L. Talbot. 2013. Development and characterization of 21 polymorphic microsatellite markers for the barren-ground shrew, Sorex ugyunak (Mammalia: Sorcidae), through next-generation sequencing, and cross-species amplification in the masked shrew, Sorex cinereus. Conservation Genetics Resources 5(2):315-318. doi:10.1007/s12686-012-9792-5 [Details] [Full Publication]

Yezerinac, S., R. B. Lanctot, S. L. Talbot, and G. K. Sage. 2013. Social and genetic mating system of American Golden-Plovers. Condor 115(4):808-815. doi:10.1525/cond.2013.120081 [Details] [Full Publication]

Campbell, M. A., G. K. Sage, J. A. López, R. L. DeWilde, and S. L. Talbot. 2013. Development and characterization of 16 polymorphic microsatellite loci for the Alaska blackfish (Esociformes: Esocidae: Dallia pectoralis). Conservation Genetics 6(2):349-351. doi:10.1007/s12686-013-0091-6 [Details] [Full Publication]

Roffler, G. H., L. G. Adams, S. L. Talbot, G. K. Sage, and B. W. Dale. 2012. Range overlap and individual movements during breeding season influence genetic relationships of caribou herds in southcentral Alaska. Journal of Mammalogy 93(5):1318-1330. doi:10.1644/11-MAMM-A-275.1 [Details] [Full Publication]

Sonsthagen, S. A., T. J. Coonan, B. C. Latta, G. K. Sage, and S. L. Talbot. 2012. Effect of a recent colonization on genetic diversity of a territorial raptor the Golden Eagle. Biological Conservation 146(1):116-122. doi:10.1016/j.biocon.2011.11.031 [Details] [Full Publication]

Sonsthagen, S. A., E. L. McClaren, F. I. Doyle, K. Titus, G. K. Sage, R. E. Wilson, J. R. Gust, and S. L. Talbot. 2012. Identification of metapopulation dynamics in Northern Goshawk of the Alexander Archipelago and coastal British Columbia. Conservation Genetics 13(4):1045-1057. doi:10.1007/s10592-012-0352-z [Details] [Full Publication]

Miller, W., S. C. Schuster, A. J. Welch, A. Ratan, O. C. Bedoya-Reina, F. Zhao, H. L. Kim, R. C. Burhans, D. I. Drautz, N. E. Wittenkindt, L. P. Tomsho, E. Ibarra-Laclette, L. Herrera-Estrella, E. Peacock, S. D. Farley, G. K. Sage, K. D. Rode, M. E. Obbard, R. Montiel, L. Bachmann, Ó. Ingólfsson, J. Aars, T. Mailund, Ø. Wiig, S. L. Talbot, and C. Lindqvist. 2012. Polar and brown bear genomes reveal ancient admixture and demographic footprints of past climate change. Proceedings of the National Academy of Sciences of USA 109:14295-14296. doi:10.1073/pnas.1210506109 [Details] [Full Publication]

Sonsthagen, S. A., C. V. Jay, A. S. Fischbach, G. K. Sage, and S. L. Talbot. 2012. Spatial genetic structure and asymmetrical gene flow within the Pacific walrus. Journal of Mammalogy 93(6):1512-1524. doi:10.1644/11-MAMM-A-344.1 [Details] [Full Publication]

Dial, C. R., S. L. Talbot, G. K. Sage, M. Seidensticker, and D. W. Holt. 2012. Cross-species amplification of microsatellite markers in the Great Horned Owl Bubo virginianus, Short-eared Owl Asio flammeus and Snowy Owl B. scandiacus for use in population genetics, individual identification and parentage studies. Yamashina Institute for Ornithology 44:1-12. [Details]

Gangoso, L., M. Alcaide, J. M. Grande, J. Muñoz, S. L. Talbot, S. A. Sonsthagen, G. K. Sage, and J. Figuerola. 2012. Colonizing the world in spite of reduced MHC variation. Journal of Environmental Biology 25(7):1438-1447. doi:10.1111/j.1420-9101.2012.02529.x [Details] [Full Publication]

Williams, I., B. M. Guzzetti, J. R. Gust, G. K. Sage, R. E. Gill, Jr, T. L. Tibbitts, S. A. Sonsthagen, and S. L. Talbot. 2012. Polymorphic microsatellite loci identified through development and cross-species amplification within shorebirds (Charadriidae and Scolopacidae). Journal of Ornithology 153(2):593-601. doi:10.1007/s10336-011-0811-1 [Details] [Full Publication]

Toussaint, R. K., D. Scheel, G. K. Sage, and S. L. Talbot. 2012. Nuclear and mitochondrial markers reveal evidence for genetically segregated cryptic speciation in giant Pacific octopuses from Prince William Sound, Alaska. Conservation Genetics 13:1483-1497. doi:10.1007/s10592-012-0392-4 [Details] [Full Publication]

Booms, T. L., S. L. Talbot, G. K. Sage, B. J. McCaffery, K. G. McCracken, and P. F. Schempf. 2011. Nest-site fidelity and dispersal of gyrfalcons estimated by noninvasive genetic sampling. Condor 113(4):768-778. doi:10.1525/cond.2011.100178 [Details] [Full Publication]

Talbot, S. L., A. Palmer, G. K. Sage, S. A. Sonsthagen, T. Swem, D. Brimm, and C. M. White. 2011. Lack of genetic polymorphism among Peregrine Falcons of Fiji. Journal of Avian Biology 42:415-428. [Details]

Toussaint, R. K., G. K. Sage, S. L. Talbot, and D. Scheel. 2011. Microsatellite marker isolation and development for the giant Pacific octopus (Enteroctopus dofleini). Conservation Genetics Resources 4(3):545-548. doi:10.1007/s12686-011-9588-z [Details] [Full Publication]

Johnson, J. A., S. L. Talbot, G. K. Sage, K. K. Burnham, J. W. Brown, T. L. Maechtle, W. S. Seegar, M. A. Yates, B. Anderson, and D. P. Mindell. 2010. The Use of Genetics for the Management of a Recovering Population: Temporal Assessment of Migratory Peregrine Falcons in North America. PLoS One 5(11):e14042. doi:10.1371/journal.pone.0014042 [Details] [Full Publication]

Cronin, M. A., S. C. Amstrup, S. L. Talbot, G. K. Sage, and K. S. Amstrup. 2009. Genetic variation, relatedness, and effective population size of polar bears (Ursus maritimus) in the southern Beaufort Sea, Alaska. Journal of Heredity 100:681-690. [Details]

Fischbach, A. S., C. V. Jay, J. V. Jackson, L. W. Andersen, G. K. Sage, and S. L. Talbot. 2008. Molecular method for determining sex of walruses. Journal of Wildlife Management 72(8):1808-1812. doi:10.2193/2007-413 [Details] [Full Publication]

Roffler, G. H., L. G. Adams, G. Luikart, G. K. Sage, R. Schwanke, M. K. Schwartz, S. L. Talbot, and K. L. Pilgrim. 2008. Development of a technique to extract DNA from Dall's sheep feces. Proceedings: North American Wild Sheep and Goat Council 16th Biennial Symposium, April 27-May 1, 2008. [Details]

Ramstad, K. M., C. A. Woody, C. Habicht, G. K. Sage, J. E. Seeb, and F. W. Allendorf. 2007. Concordance of nuclear and mitochondrial DNA markers in detecting a founder event in Lake Clark sockeye salmon. American Fisheries Society Symposium 54:31-50. [Details] [Full Publication]

Muñiz-Salazar, R., S. L. Talbot, G. K. Sage, D. H. Ward, and A. Cabello-Pasini. 2006. Genetic structure of eelgrass Zostera marina meadows in an embayment with restricted water flow. Marine Ecology Progress Series 309:107-116. [Details] [Full Publication]

Talbot, S. L., S. Wyllie-Echeverria, D. H. Ward, J. R. Rearick, G. K. Sage, B. Chesney, and R. C. Phillips. 2006. Genetic characterization of Zostera asiatica on the Pacific Coast of North America. Aquatic Botany 85(3):169-176. doi:10.1016/j.aquabot.2006.03.011 [Details] [Full Publication]

Handel, C. M., L. M. Pajot, S. L. Talbot, and G. K. Sage. 2006. Use of buccal swabs for sampling DNA from nestling and adult birds. Wildlife Society Bulletin 34(4):1094-1100. doi:10.2193/0091-7648(2006)34[1094%3AUOBSFS]2.0.CO%3B2 [Details] [Full Publication]

Roffler, G. H., L. G. Adams, S. L. Talbot, and G. K. Sage. 2006. Evaluating the genetic relationships of the Mentasta and Nelchina caribou herds in Southcentral Alaska. Proceedings: 11th North American Caribou Conference, 23-27 April, 2006. doi:10.7557/ [Details] [Full Publication]

Weckworth, B. V., S. L. Talbot, G. K. Sage, D. K. Person, and J. A. Cook. 2005. A signal for independent coastal and continental histories among North American wolves. Molecular Ecology 14(4):917-931. doi:10.1111/j.1365-294X.2005.02461.x [Details] [Full Publication]

Muñiz-Salazar, R., S. L. Talbot, G. K. Sage, D. H. Ward, and A. Cabello-Pasini. 2005. Population genetic structure of annual and perennial populations of Zostera marina L. along the Pacific coast of Baja California and the Gulf of California. Molecular Ecology 14(3):711-722. doi:10.1111/j.1365-294X.2005.02454.x [Details] [Full Publication]

Talbot, S. L., J. R. Gust, G. K. Sage, A. S. Fischbach, K. S. Amstrup, W. Leacock, and L. J. Van Daele. 2005. Genetic characterization of brown bears of the Kodiak Archipelago. Final Report to the U.S. Fish and Wildlife Service, Kodiak National Wildlife Refuge, USGS Alaska Science Center, Anchorage, Alaska, 87 p. [Details]

Ramstad, K. M., C. A. Woody, G. K. Sage, and F. W. Allendorf. 2004. Founding events influence genetic population structure of sockeye salmon (Oncorhynchus nerka) in Lake Clark, Alaska. Molecular Ecology 13:277-290. doi:10.1046/j.1365-294X.2003.2062.x [Details] [Full Publication]

Talbot, S. L., J. R. Gust, G. K. Sage, R. E. Gill, Jr, T. L. Tibbitts, R. B. Lanctot, and B. J. McCaffery. 2004. Population differentiation of breeding Bristle-thighed Curlews (Numenius tahitiensis). USGS Unpublished Report. [Details]

Woody, C. A., K. M. Ramstad, D. B. Young, G. K. Sage, and F. W. Allendorf. 2004. Lake Clark sockeye salmon population assessment. Final Report to the U.S. Fish and Wildlife Service Federal Office of Subsistence Management, Fisheries Resource Monitoring Program (FIS 01-042). [Details]

Olsen, J. B., W. J. Spearman, G. K. Sage, S. J. Miller, B. G. Flannery, and J. K. Wenburg. 2004. Variation in the population structure of Yukon River chum and coho salmon: Evaluating the potential impact of localized habitat degradation. Transactions of the American Fisheries Society 133:476-483. doi:10.1577/03-041 [Details] [Full Publication]

Nielsen, J. L., I. Williams, G. K. Sage, and C. E. Zimmerman. 2003. The importance of genetic verification for determination of Atlantic salmon in north Pacific waters. Journal of Fish Biology 62(4):871-878. doi:10.1046/j.1095-8649.2003.00072.x [Details] [Full Publication]

Nielsen, J. L., G. K. Sage, S. A. Pavey, R. Valenzuela, and T. C. Wiacek. 2003. Hatchery and wild steelhead genetics in Idaho: Report #1: Dworshak Hatchery stocks in the Clearwater drainage. Technical Report submitted to Idaho Department of Fish and Game, April 2003. [Details]

Scribner, K. T., G. K. Sage, C. Soiseth, L. K. Thorsteinson, E. E. Knudsen, and J. L. Nielsen. 2003. Genetic analysis of coho salmon colonization in recently deglaciated streams in Glacier Bay, Alaska. Report submitted to National Park Service, USGS Alaska Science Center, Anchorage, Alaska. [Details] [Full Publication]

Nielsen, J. L. and G. K. Sage. 2002. Microsatellite sequence reveals significant haplotype diversity in Central Valley chinook stocks. California Dept. of Fish and Game, June 2002. [Details]

Nielsen, J. L. and G. K. Sage. 2002. Genetic analysis of temporal spawning populations of Lahontan cutthroat (Oncorhynchus clarki henshawi) from Pyramid Lake. Bailey Environmental Consulting, 11 p. [Details]

Nielsen, J. L. and G. K. Sage. 2002. Population genetic structure in Lahontan cutthroat trout. Transactions of the American Fisheries Society 131:376-388. [Details]

Nielsen, J. L. and G. K. Sage. 2002. Microsatellite analyses of San Mateo Creek trout, 2001. Technical Report submitted to U.S. Department of Defense, Camp Pendleton, CA. [Details]

Nielsen, J. L. and G. K. Sage. 2001. Microsatellite analyses of the trout of Northwest Mexico. Genetica 111(No.1-3):269-278. [Details] [Full Publication]

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